PUBLICACIONES

Rodríguez Esquivel, S. V., Quesada-Perez, F., Nova-Bustos, N., García-Rojas, A. and Alvarado, J. .J. (2025). Prevalence of Diseases in Scleractinian Corals of the Southern Caribbean of Costa Rica. Caribbean Journal of Science, 55(1):156-175. https://doi.org/10.18475/cjos.v55i1.a15 

Hidalgo-Sánchez, R., Alvarado, J.J., Araya-Valverde, E., and  Díaz-Ferguson, E. (2025). Genetic diversity and structure of Pocillopora spp. corals in the Gulf of Papagayo, Costa Rica. Discov Oceans 2, 16. https://doi.org/10.1007/s44289-025-00055-8

Montiel-Barrantes, M. P., Brenes-Guillén, L., Carvajal-Rojas, S. ., González-Sánchez, K. ., Cordón-Krumme, M. I. ., & Alvarado, J. J. (2025). Connectivity of the sea urchin Diadema mexicanum A. Agassiz, 1863 (Echinoidea, Diadematidae) in the Pacific coast of Costa Rica. Revista Ciencias Marinas Y Costeras, 17(1), 91-115. https://doi.org/10.15359/revmar.17-1.5

 Fabregat-Malé, S., & Alvarado, J. J. (2025). A story of disturbance and loss: historical coral reef degradation in Bahía Culebra, North Pacific of Costa Rica. Revista De Biología Tropical, 73(S1), e63624. https://doi.org/10.15517/rev.biol.trop.v73iS1.63624 

 Alvarado, J. J., Evans, K., Kleypas, J. A., Marín-Moraga, J. A., Mendez-Venegas, M., Pérez-Reyes, C., Sandoval, M., Solano, M. J., & Villalobos-Cubero, T. (2025). Coral reefs restoration initiatives in Costa Rica: ten years building hope. Revista De Biología Tropical, 73(S1), e63695. https://doi.org/10.15517/rev.biol.trop.v73iS1.63695 

Mena, S., Quesada-Perez, F., Sánchez-Noguera, C., Salas-Moya, C., Alvarado, J. J., López-Garro, A., Zanella, I., Méndez Venegas, M., Piedra-Chacón, R., Anderson Rana, L., & Fonseca López, L. (2025). Community Structure of coral ecosystems in sites of importance for the conservation of marine biodiversity in the North Pacific of Costa Rica . Revista De Biología Tropical, 73(S1), e63715. https://doi.org/10.15517/rev.biol.trop.v73iS1.63715 

 Gutierrez, L., Polidoro, B., Obura, D., Cabada-Blanco, F., Pettersson, E., Pearce-Kelly, P., Kemppinen, K., Linardich, C., Alvarado, J.J., Alvarez-Filip, L., Banaszak, A., Casado de Amezua, P., Crabbe, J., Croquer, A., Feingold, J., Goergen, E., Goffredo, S., Hoeksema, B., Huang, D., Kennedy, E., Kitahara, M., Kružić, P.,  Miller, M., Nunes, F., Quimbayo, J.P., Rivera-Sosa, A., Rodríguez-Martínez, R., Santodomingo, N., Sweet, M., Vermeij, M., Villamizar, E., Aeby, G.,  Alliji, A., Couce, E., Cowburn, B., Porter, S., Shlesinger, T., Bayley, B., Wilson, B., Nuñez Lendo, C.I., Samimi-Namin, K., & Kersting, D. 2024. Half of Atlantic reef-building corals at elevated risk of extinction due to climate change and other threats. PLOS ONE 19(11): e0309354. https://doi.org/10.1371/journal.pone.0309354  

 Zúñiga-Madrigal, J., Angulo-Vázquez, J., López-García, Y., Barrantes, G., & Alvarado, J.J. 2024. Guardianes del Ecosistema: herramienta lúdico-didáctica para fomentar la conservación de ecosistemas vulnerables en estudiantes de primaria en Costa Rica. Revista Latinoamericana de Ciencias Sociales y Humanidades V (5); 2799. https://latam.redilat.org/index.php/lt/article/view/2815/4373 

Fabregat-Malé, S., Mena-González, S., Quesada-Perez, F., & Alvarado, J.J. 2024. Testing the feasibility of coral nurseries in an upwelling area in the North Pacific of Costa Rica. Front. Mar. Sci. 11:1400026. https://doi.org/10.3389/fmars.2024.1400026 

 Bogantes-Retana, A., Barquero-Jackson, J.,  Vargas-Guerrero, M., & Alvarado, J.J. (2024).  Variation of the diet of the sea urchin (Diadema mexicanum (Diadematoida: Diadematidae)) according to its size in the Eastern Tropical Pacific. Revista de Biología Tropica, 72(S1), e58602.  https://doi.org/10.15517/rev.biol.trop..v72iS1.58602 

Hernández, J. C., Alfonso, B., Guzmán-Mora, A. G., & Alvarado, J. J. (2024). Predators of the sea urchin Diadema mexicanum (Diadematoida: Diadematidae) at the Eastern Tropical Pacific coral reefs. Revista De Biología Tropical, 72(S1), e59007. https://doi.org/10.15517/rev.biol.trop.v72iS1.59007 

Uthicke, S., Pratchett, M.S., Bronstein, O., Alvarado, J.J., & Wörheide, G. (2024) The crown‑of‑thorns seastar species complex: knowledge on the biology and ecology of five corallivorous Acanthaster species. Marine Biology 171:32. https://doi.org/10.1007/s00227-023-04355-5   

May-Collado, L.J., Bottoms, S., Durant, G., Palacios-Alfaro, J.D., & Alvarado, J.J. (2023). Southern Hemisphere humpback whales (Megaptera novaeangliae) (Artiodactyla: Balaenopteridae) singing activity at Caño Island Biological Reserve, Costa Rica before, during, and after COVID-19 lockdowns. Revista de Biología Tropical Vol. 71(S4): e57280. https://doi.org/10.15517/rev.biol.trop..v71iS4.57280

Quezada-Perez F, Mena S, Fernández-García C, Alvarado JJ. (2023). Status of Coral Reef Communities on the Caribbean Coast of Costa Rica: Are We Talking about Corals or Macroalgae Reefs? Oceans 4(3):315-332. https://doi.org/10.3390/oceans4030022 

Dubuc, A., Quimbayo, J.P., Alvarado, J.J., Araya-Arce, T., Arriaga, A., Ayala-Bocos, A., Casas-Maldonado, J.J, Chasqui, L., Cortés, J., Cupul-Magaña, A., Olivier, D., Olán-González, M., González-Leiva, A., Reyes-Bonilla, H., Smith, F., Rivera, F., Rodríguez-Zaragoza, F.A., Rodríguez-Villalobos, J.C.,  Segovia, J., Zapata, F.A., & Bejarano, S. (2023). Patterns of reef fish taxonomic and functional diversity in the Eastern Tropical  Pacific. Ecography 2023: e06536 https://doi.org/10.1111/ecog.06536 

Chomitz, B.R., Kleypas, J.A., Cortés, J., & Alvarado, J.J. (2023). Change in the composition of fauna associated with Pocillopora spp. (Scleractinia, Pocilloporidae) following transplantation. Revista de Biología Tropical 71 (S1): e54882. https://doi.org/10.15517/rev.biol.trop..v71iS1.54882   

Palou-Zuñiga, N., Madrigal-Ballestero, R., Schlüter, J., & Alvarado, J.J. (2023). Applying the SES framework to coral restoration projects on the Pacific coast of Costa Rica. Revista de Biología Tropical 71 (S1): e54853. https://doi.org/10.15517/rev.biol.trop..v71iS1.54853 

Fabregat-Malé, S., Mena, S., & Alvarado, J.J. (2023). Nursery-reared coral outplanting succes in an upwelling-influenced area in Costa Rica. Revista de Biología Tropical 71 (S1): e54879. https://doi.org/10.15517/rev.biol.trop..v71iS1.54879 

Chomitz, B.R., Kleypas, J.A., Cortés, J., & Alvarado, J.J. (2023). Succesion of sessile benthic community at a coral reef restoration site. Revista de Biología Tropical 71 (S1): e54881. https://doi.org/10.15517/rev.biol.trop..v71iS1.54881 

Combillet, L., Fabregat-Malé, S., Mena, S., Marín-Moraga, J.A., Gutiérrez, M., & Alvarado, J.J. (2022). Pocillopora spp. growth analysis on restoration structures in an Eastern Tropical Pacific upwelling area. PeerJ 10:e1 3248 https://doi.org/10.7717/peerj.13248 

Alvarado, J.J., Azofeifa-Solano, J.C., Chacón-Monge, L. & Cortés, J. (2022) Diversity of deep-water echinoderms from Costa Rica. Frontiers In Marine Science. https://doi.org/10.3389/fmars.2022.918878

Arias-Godínez, G.A., Jimenez, C., Gamboa, C., Cortés, J., Espinoza, M., & Alvarado, J.J. (2021). The effect of coral reef degradation on the trophic structure of reef fishes from Bahía Culebra, north Pacific coast of Costa Rica. Journal of Coastal Conservation 25, 8 (2021). https://doi.org/10.1007/s11852-021-00802-x

Salas-Moya, C., R. Vargas-Castillo, JJ Alvarado, J.C Azofeifa-Solano & J. Cortés. (2021). Decapods associated with macroinvertebrates in the Pacific of Costa Rica. Marine Biodiversity Records 14:6. https://doi.org/10.1186/s41200-020-00199-w

Chacón-Monge, J.L., Azofeifa-Solano, J.C., Alvarado, J.J., & Cortés, J. (2021). Área de Conservación Guanacaste Echinoderms, North Pacific of Costa Rica. Revista de Biología Tropical, 69(S1), 487-500. https://doi.org/10.15517/RBT.V69ISUPPL.%201.46391

Salas-Moya, C., Fabregat-Malé, S., Vargas-Castillo, R., Valverde, J.M., Vásquez-Fallas, F., Sibaja-Cordero, J., Alvarado, J.J. (2021) Pocillopora cryptofauna and their response to host coral mortality. Symbiosis 84:91–103. https://doi.org/10.1007/s13199-021-00771-7

Fernández-García, C., Salas-Moya, C., Mena, S., Azofeifa-Solano, J.C., & Alvarado, J.J. (2021) Diversidad de los hábitats submareales de la Península de Santa Elena e Islas Murciélago, Pacífico Norte, Costa Rica. Revista de Biología Tropical 69 (Suppl. 2): S160-S179. https://doi.org/10.15517/rbt.v69iS2.48774 

González-Sánchez, K., Flores-Alvarado, B., Montiel-Barrantes, P., Gómez-Arce, G., & Alvarado, J.J. (2021). Ascidian diversity of Costa Rica, including new records for the North Pacific. Revista de Biología Tropical 69(Suppl. 2): S234-S245. https://doi.org/10.15517/rbt.v69iS2.48318.

Alvarado, J. J. & Fabergat-Malé, S. (2021). Echinoderm research perspectives: A Central American bibliometric review. UNED Research Journal, 13(2), e3535. DOI: 10.22458/urj.v13i2.3535 

Alvarado, J. J., Azofeifa-Solano, J. C., Beita-Jiménez, A., Cortés, J., Mena, S., Salas-Moya, C., & Fernández-García, C. (2021). Reef environments of Murciélago Islands and Santa Elena Peninsula, Guanacaste Conservation Area, Costa Rican Pacific. Scientia Insularum. https://doi.org/10.25145/j.SI.2021.04.004

Fernández-Rivera F.J., Espino-Barrb E., López–Rocha, JA., Reyes-Bonilla H, Alvarado J.J. (2020). Growth parameters and natural mortality of Diadema mexicanum A. Agassiz, 1863 in the coast of Colima, México. Cahiers de Biologie Marine 61:459-467. https://doi.org/10.21411/CBM.A.4D7E8748

Solís-Marín, F.A., Alvarado, J.J., Conejeros Vargas, C.A., & Caballero Ochoa, A.A. (2020). Pentamera fonsecae sp. nov. a new species of sea cucumber from the Pacific coast of Costa Rica (Holothuroidea, Dendrochirotida, Thyonidae). Zootaxa 4878 (3): 581–588. https://doi.org/10.11646/zootaxa.4878.3.9

Quimbayo, J.P., Dias, M.S., Kulbicki, M., Mendes, T.C., Lamb, R.W., Johnson, A.F., Aburto-Oropeza, O., Alvarado, J.J., Bocos, A.A., Ferreira, C.E.L., Garcia, E., Luiz, O.J., Mascareñas-Osorio, I., Pinheiro, H.T., Rodriguez-Zaragoza, F., Salas, E., Zapata, F.A. & Floeter, S.R. (2019). Determinants of reef fish assemblages in tropical oceanic islands. Ecography, 42 (1): 77–87. https://doi.org/10.1111/ecog.03506

Beita-Jiménez, A., J.J. Alvarado, S. Mena, & A.G. Guzmán-Mora. (2019) Benefits of protection on reef fish assemblages in a human impacted region in Costa Rica. Ocean and Costal Management, 169, 165-170. https://doi.org/10.1016/j.ocecoaman.2018.12.023

Auster P.J., Cortés, J., Alvarado, J.J., & and Beita-Jiménez, A. (2019). Coordinated hunting behaviors of mixed-species groups of piscivores and associated species at Isla del Coco National Park (Eastern Tropical Pacific). Neotropical Ichthyology, 17(1) Epub11-Mar-2019. doi.org/10.1590/1982-0224-20180165

Fourriére, M., Alvarado, J.J., Cortés, J., Taylor, M., Ayala-Bocos, A., Heidemeyer, M., Arauz, R., López-Garro, A., & Wolff, M. (2019). Energy flow structure and role of keystone groups in shallow water environments in Isla del Coco, Costa Rica, Eastern Tropical Pacific. Ecological Modelling, 396; 74-85. https://doi.org/ 10.1016/j.ecolmodel.2019.01.004

Arias-Godínez, G., Jiménez, C., Gamboa, C., Cortés, J., Espinoza, M., & Alvarado, J.J. (2019). Spatial and temporal changes of reef fish assemblages on disturbed coral reefs, north Pacific coast of Costa Rica. Marine Ecology: an evolutionary perspective, 40: e12532. https://doi.org/10.1111/maec.12532

Cambronero-Solano, S. Benavides, R., Solís-Marín, F. A., & Alvarado, J.J. (2019) New reports of echinoderms on the Caribbean Continental Shelf of Central America. Zoosymposia doi.org/10.11646/zoosymposia.0.0.0

Alvarado, J.J., C. Sánchez-Noguera, G. Arias-Godínez, T. Araya, C. Fernández-García, & A.G. Guzmán. (2019). Impact of El Niño 2015-2016 on the coral reefs of the Pacific of Costa Rica: the potential role of marine protection. Revista de Biología Tropical 8(Suppl. 1), S271-S282. https://doi.org/10.15517/rbt.v68iS1.41190

Alvarado, J.J., A. Beita-Jiménez, S. Mena, C. Fernández-García, J. Cortés, C. Sánchez-Noguera, C. Jiménez, & A.G. Guzmán. (2018). Cuando la conservación no puede seguir el ritmo del desarrollo: Estado de salud de los ecosistemas coralinos del Pacífico Norte de Costa Rica. Revista de Biología Tropical 66 (Suppl. 1). 280-308. https://doi.org/10.15517/rbt.v66i1.33300

Azofeifa-Solano, J.C, S. Mena, J.J. Alvarado, J. L. Chacón-Monge, T.M. Clarke, J. Herrera-Correal and I.S. Wehrtmann. (2017). Echinoderms from a near-estuarine system in the Costa Rican Pacific, Central America. Check List 13: 2113. https://doi.org/10.15560/13.3.2113

Alvarado, J.J., J. L. Chacón-Monge, F.A. Solís-Marín, T. Pineda-Enríquez, A.A. Caballero-Ochoa, S. Solano-Rivera & R. Romero-Chaves. (2017).Equinodermos del Museo de Zoología de la Universidad de Costa Rica. Revista de Biología Tropical 65 (Supl. 1): S272-S287. https://doi.org/

Beita-Jiménez, A., & J.J. Alvarado. (2016). Anomalous coloration of a yellow tail damselfish Stegastes arcifrons (Actinopterygii: Pomacentridae) at Isla del Coco National Park, Costa Rica. Revista Biología Tropical 64 (Supl. 1):197-199. https://doi.org/10.15517/rbt.v64i1.23447

Alvarado, J.J., A. Beita, S. Men, C. Fernández-García, A.G. Guzmán, & J. Cortés. (2016). Ecosistemas coralinos del Parque Nacional Isla del Coco, Costa Rica: análisis estructural y temporal. Revista Biología Tropical 64 (Supl. 1): 153-175.https://doi.org/10.15517/rbt.v64i1.23423

Alvarado JJ, J. Cortés, H.M. Guzman & H. Reyes-Bonilla. 2016. Distribution, size and diet of Diadema mexicanum (Echinoidea) along the Eastern Tropical Pacific coral reefs. Aquatic Biology 24: 151-161. https://doi.org/10.3354/ab00645

Paz-García, D.A., C. Galván-Tirado, J.J. Alvarado, Jorge Cortés, F. J. García-De-Leó, M.E. Hellberg, & E.F. Balart. (2016). Variation in the whole mitogenome of reef-building Porites corals. Conservation Genet Resour 8(2):123-127. https://doi.org/10.1007/s12686-016-0527-x

Alvarado JJ, J. Cortés, H.M. Guzman & H. Reyes-Bonilla. (2016).  Bioerosion by the sea urchin Diadema mexicanum along Eastern Tropical Pacific coral reef. Marine Ecology: an evolutionary perspective. 37(5): https://doi.org/10.1111/maec.12372

Fourriére, M., JJ Alvarado, J Cortés & A. Ayala-Bocos. (2016). Updated checklist and analysis of completeness of the marine fish fauna of Isla del Coco, Costa Rica. Marine Biodiversity 47(3): https://doi.org/10.1007/s12526-016-0501-6

Alvarado, JJ., A. Beita, S. Mena, C. Fernández-García & A.G. Guzmán. (2015). Ecosistemas coralinos del Área de Conservación Osa, Costa Rica: análisis estructural y necesidades de conservación. Revista de Biología Tropical 63 (Supl. 1):219-259. https://doi.org/10.15517/rbt.v63i1.23105

Alvarado, J.J., H. Reyes-Bonilla & F. Benítez-Villalobos. (2015). Diadema mexicanum, erizo der mar clave en los arrecifes coralinos del Pacifico Tropical Oriental: estado de conocimiento y perspectivas futuras. Revista Biología Tropical 63 (Supl. 2): 135-157. https://doi.org/10.15517/rbt.v63i2.23140